Exoprosopini  

The members of the Australian Exoprosopini can be distinguished from other Australian Anthracinae by the antennae lacking an apical circlet of hairs, usually with a well developed basal stylomere; thoracic laterotergites and mediotergites haired, plumula and hairs upon tympanal ridge well developed; sharp, long basicosta at base of costal vein, R2+3 arising from R with distinct basal 90˚ bend at r-m cross-vein, patagium of scales present at base of wing, squama and alula fringed with scales; reduced conical pulvilli that appear as basal teeth to tarsal claws; males with aedeagus enclosed within epiphallus, emerging through deep notch on medial ventral surface; females with a sand chamber formed from a sclerotised sheet-like S8 producing a large, bilobed, hemispherical, posteriorly open cavity; T8 is invaginated, with a dorsal apodeme or distinct marginal thickening, abruptly reduced laterally to long, narrow, apically-directed flanges, posterior brush of hairs are sparse and reduced; furca a single U-shaped sclerite, strong broad arms apically inturned to form dorsal hook, basal sclerite broad sculptured. Ten genera represent the tribe in Australia; Atrichochira, Balaana gen. nov., Exoprosopa, Kapua, Ligyra, Larrpana, Munjua, Muwarna, Palirika, Pseudopenthes, and Wurda.

Comments. The Villini also has a patagium of scales, sharp basicosta, R2+3 arising from R with a basal 90˚ bend at i-r cross-vein, females with lateral flanges to T8, sheet-like S8, and U-shaped furca. In the Villini the aedeagus is not enclosed within the epiphallus, but distinctly separate from the base, and the pulvilli are not smooth glabrous cones.